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In the early 1990s, the parcels sector of British Rail was seeking a fleet of multiple units that would be more cost-effective to run than the locomotive hauled stock it was using. The initial proposal was to convert a number of redundant Class 307 units that were in the process of being withdrawn from passenger service into Class 300 parcels units. Owing to the age of these units however, it was subsequently decided to procure a fleet of new trains. These units, which were ordered in 1994, were initially given the TOPS classification Class 350, before being changed to Class 325.

The 16 units were built at ABB Derby between 1995 and 1996. They are very similar to Class 319 units, sharing the same traction equipment and body design, but are fitted with cabs of the same design as the ABB Networker family.

TOPS numbers are on the front of the cabs under the driver's window in a non-standard typeface (Plantin, the then corporate typeface of Royal Mail), and the units were numbered 325001 - 325016. The units are fitted with large round oleo buffers, and have no gangways between carriages. They carry a livery of Royal Mail red, with two yellow stripes running along the lower bodyside before turning sharply backwards and pointing up towards the roof, black cab window surrounds and a full yellow warning panel. Each set is made up of four cars, with roller doors in place of sliding ones and no windows. Each car has two roller shutter sliding doors on each side and is designed to hold up to 12 tonnes. They have a pantograph to pick up power from the 25 kV AC overhead lines, and also a shoe to pick up power off the 750 V DC third rail. They cannot work in multiple with any other multiple unit stock, but are fitted with drop-head buck-eye coupling and can therefore be hauled by locomotives. The units were built in such a way that they could easily be converted for passenger use if no longer required for mail services.

After introduction, which was relatively trouble-free due largely to the fact that the Class 325 shares traction equipment with the Class 319, the units successfully settled into carrying parcels and mail from London to Glasgow and Edinburgh at 100 mph (160 km/h). They are based at Crewe Electric TMD.

As of 2017, the mockup cab built before the construction of the Class 325s, numbered 325000, is currently on display at the Nene Valley Railway as part of the as-yet-unfinished Night Mail Museum at Overton, having been disposed of from the National Railway Museum.